When Bengals grow up into adults, there can be some friction as each tries to exert their influence on the other and there may be fighting for territory. I find cats who are equal in status have more problems than those where there is a natural leader and a natural follower. Though a highly territorial cat can start bullying a weaker one too. I think the basement incident, catapulted both into reassessing their status in the household, and so now you have ongoing issues. I guess the day that the younger one had the run of the house, left him with the feeling the house was HIS territory and the other one is now the interloper into HIS space. He probably thought the older one had naturally gone to find pastures new as they would do in the wild. I think I would try and allow them to have separate territories, so that they do not have to meet too often, duplicate the food and the litter trays and place them in each "territory". Introduce some vertical space or more vertical space if you already have some. Cats are territorial, Bengals I believe are often very territorial and I guess with or without the basement incident you could possibly be facing the same issue. On a better note, I had this problem at around the same age with two males who grew up together but suddenly there was serious fighting. Rolling on the floor, fur everywhere stuff, they couldn't stand each other and I separated them, never let them meet unattended. But years later they calmed down and became best buddies again. The other strategy that I have heard of, is to lock them both in a small space with separate places to hide and leave them to it for a few days/weeks, they will sort themselves out, but that could result in injury I guess.
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